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Taxonomic Study of the Genus Hatchiana in Japan (Coleoptera, Cantharidae, Podabrini)

Shoichi IMASAKA
Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, 812-8581 Japan

Abstract.  The genus Hatchiana Fender is redefined, and all 12 Japanese species are revised and newly classified into 3 species-groups. Three new species, Hatchiana unzenensis, H. nishidai and H. sanoi are described from Japan. Nine species are newly transferred to the genus Hatchiana from the genus Podabrus and redescribed. Keys to the species-groups and species in Japan are provided. The 1st to 3rd antennal segments, femur of each leg, female 8th abdominal sternite and male genitalia are illustrated and a distribution map is given for each species.

*Contribution from the Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka (Ser. 5, No. 69).

The genus Hatchiana was proposed by Fender (1966) for H. arizonensis based on a single male from Arizona, North America and was admitted as an independent genus in Podabrini by Delkeskamp (1977). Later Kasantsev (1992) treated Hatchiana as a subgenus in the genus Podabrus using the features of male genitalia and claws.
According to the synopsis of the Telephorinae (Westwood, 1840), the genus Podabrus was created by Fischer de Waldheim in 1821 based on Telephorus alpinus Paykull. However, the original description by Fischer de Waldheim (1821) has never been found in any of Fischer's published papers (Lacordaire, 1857). Thereafter the author of Podabrus has been attributed to Westwood 1840 according to the ICZN, and the Fischer's generic name was treated as nomen nudum in Neaves (1940).
In Japan, at least 90 species have been placed in the genus Podabrus. By examining all these species, they are possibly divided into 4 species groups, of which 1 includes 9 species described by Kiesenwetter (1879), Makino & Nakane (1982), Sato (1986), Takakura (1988), Takahashi (1992) and Kiriyama (2000) based on appearance, body coloration, male genitalia, etc. These 9 species have many important features common to those of Hatchiana. However, the features characterizing Hatchiana in the original description (Fender, 1966) includes those specialized to the type species, since the genus was created based on a single male.
In this paper, the aforementioned 9 species are redescribed and their generic position is examined by compareing their morphological features with those of Hatchiana and other related subgenera of the genus Podabrus. At the same time, a taxonomic status of Hatchiana is determined and its range of morphological features is redefined. In addition, 3 new species are described and Hatchiana is divided into 3 species -groups. Keys to the species-groups and species in Japan are also provided.
Taxonomic status of Hatchiana
The genus Hatchiana was originally defined based on the following features (Fender, 1966): (1) apical segment of antennae distorted; (2) membranous lobe of maxillae produced; (3) male genitalia with an additional pair of conical, hirsute, sclerotized processes between dorsal plate and penical lobe; the processes extending beyond dorsal plate and connected to the base of penial lobe by long, slender filaments; (4) dorsal plate of paramere short, truncate at apex, with a small median notch; (5) pro- and mesotarsal claws broadly cleft; claw apically divided into 2 portions, which are nearly parallel in pro- and mesotarsal claws, curved and diverging from the claw in metatarsal claw.
Among these features, (1), (2) and (3) are restricted to the type species and not found in the 9 Japanese species. Many other characters including (4) and (5) are shared by the type species and the Japanese species. In particular, fundamental features in male genitalia indicate that they should be placed in Hatchiana as shown below in the descriptions and redescriptions of respective species.
Kasantsev (1992) treated Hatchiana as a subgenus in the genus Podabrus since both Hatchiana and Podabrus have a pair of well developed median hooks in the male genitalia and there is almost no sexual dimorphism in the shape of claws. However, Hatchiana is considered to be independent from the genus Podabrus by the following features: antennae with 2nd segement which is distinctly shorter than 3rd; claws toothed basally with triangular processes; dorsal plate of male genitalia entire, with short median notch on posterior margin. In Podabrus, 2nd antennal segment subequal to or longer than 3rd; claws toothed basally with finger-like processes; dorsal plate of male genitalia bilobed.

Genus Hatchiana Fender, 1966, stat. nov.
(Japanese name:Kubiboso-jokai-zoku)

Hatchiana Fender, 1966, Pan-Pac. Ent.., 42(4): 321-323, figs. 1-6. Type species: Hatchiana arizonensis Fender, 1966, by original designation; Delkeskamp, 1977, Col. Cat. Suppl.: 32; Brancucci, 1980, Ent. basil.., (5): 219; EIS, 1996, Nomina Ins. Nearc.1: Col., Strep.: 76;
Brachynotus Kirby, 1837 (nec De Haan, 1835). Fn. Bor. -Amer.., IV: 245-249. Type-species: Telephorus bennetii Kirby, 1837 (= Cantharis tricostatus Say, 1835), monobasic, invalid name owing to homonymy and no replaced name.
Podabrus, partim: Kiesenwetter, 1879, Dtsch. ent. Z.. 23: 306-307; Lewis, 1895, Ann. Mag. nat. Hist.., (6)16: 108; Pic, 1905, L'Echange., 21(243): 113; Makino & Nakane, 1982(1981), Rev. Fac. Sci. Kagoshima Univ. (Earth Sci. & Biol.), (14): 55-63; Sato, 1985, Col. Japan color, (III): 108; Sato, 1986, Trans. Shikoku ent. Soc.. 17(4): 255-256; Takakura, 1988, Kita-Kyushu no Konchu, 35(3): 152; Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ., 1(2): 1; Imasaka et Nishida, 1989, Koganemushi, (50): 4-5; Imasaka et al., 1990, Suzumushi, (125): 3; Takahashi, 1992, Kanagawa- chuho, (100): 75-76; Imasaka & Nakamura, 1993, Mis. Rep. Hiwa Mus. Nat. Hist., (31): 48; Imasaka & Yamaya, 1993, Bull. Nagaoka Mun. Sci. Mus., (28): 48; Takahashi, 1995, Kanagawa-chuho, (113): 20-21; Kiriyama, 1995, Keichitsu, 13(35): 1-2; Imasaka & Ohtsuka, 1996, Kumamoto Konchu Doko Kaiho, 40(3): 38.
Podabrus(Hatchiana): Kasantsev, 1992, Ent. basil..,(15): 268; Kasantsev, 1998, Ent. Rev., 76(1): 106-114, fig. 3; Takahashi & all, 1999, Bull. Kurashiki Mus. Nat. Hist., (14): 17-48; Kiriyama, 2000, Jpn. J. syst. Ent., 6(2): 249-252.
Podabrus (Podabrus): Takahashi, 1998, Kanagawa-chuho, (122): 30.
Male. Body large, 8.3-14.6mm in length; yellow to blackish brown; temples behind eyes almost black; median portion of clypeus, apical segments of labial and maxillary palpi, apical parts of mandibles, antennae from 3rd to 11th segments, disk of pronotum, meso- and metasterna, scutellum, apices of femora and tibiae, tarsi, abdomen except apical margin of each sternal segments more or less darkened (Fig. 1).
Body slender, closely covered with pubescence; apical margin of clypeus, temples behind eyes and lateral margins of pronotum fringed with bristles; elytra with bristles in addition to pubescence; lateral margins of pronotum and base of elytra smooth with faint luster.
Head strongly narrowed from eyes to occiput; anterior half of head smooth and shining, posterior half densely and rugosely punctate with a longitudinal groove at middle. Apical margin of clypeus with a pair of prominences and a subacute median notch; mandibles falcate, wide at base, narrower at apical 2/3, thin crescent in cross-section, with upper and lower margins similarly convex; eyes large, globular and strongly prominent; apical segments of labial and maxillary palpi somewhat slender, knife-shaped. Antennae slender, filiform, attaining to middle of elytra; 1st segment clavate, longer and shining; 2nd shorter than 3rd, thickened apicad with slightly luster; 3rd to 11th weakly clavate, longer and rough. Tentrium anteior in position, posterior arms of tentrium arise close to mentum at their anterior parts and auriculate, without spines from posterior bridge, gular ridges adhered to head capsule, short, rapidly close to each other behind auriculae and almost contiguous, with traces of sectoral anterior bridge at anterior margin of auriculae, dorsal arms and posterior bridge obsolete. Gular sutures double without fuse, gula between sutures strongly narrowed posteriorly behind posterior tentorial pit, then slightly dilated caudally (Fig. 3).
Pronotum subquadrate, faintly expanded backwards, wider than long; anterior margin arcuate; posterior margin weakly arcuate and bordered; lateral margins rounded; anterior angles rounded; posterior angles prominent outwards with sinus in front; disk rugose and convex at both sides near base, antero- and postro-lateral areas hollowed; medio-longitudinal furrow distinct at median area (Figs. 9, 10, 11).
Scutellum triangular with faintly rounded apex.
Elytra subparallel sided, dorsum irregularly and sparsely rugose, smooth at basal part, costate.
Legs slender; femora almost straight, sometimes ventral margin expanded at a 3rd from apex; tibiae feebly bent at base, then almost straight to apex; claws toothed, with triangular processes at base with variable shape in the following types: A type (Fig. 12) with tooth half of the length of main claw, B type (Fig. 13) with tooth 2/3 of the length of main claw, C type (Fig. 14) with tooth the same length as B type, but main claw sinuate, D type (Fig. 15) with tooth the same length as main claw, main claw sinuate, and E type (Fig. 16) with main claw elongated, tooth small and short.
Male genitalia almost oval, compact (Figs. 4-6). Tegmen embraced aedeagus (median lobe); parameres divided into dorsal and ventral lobes by deep lateral incisions (Fig. 6), dorsal lobe broad, subqudrate and subtruncate at apex, with short median notch, with setae along apical margin, with dorsal fulcrum at side angles (Fig. 4); ventral lobe narrow, tapered apically, simple, without prominence at base of lateral margins (Fig. 5). Aedeagus divided into dorsal sclerotized and ventral membranous parts (Fig. 8); dorsal sclerotized part with caudal and subcaudal sinuses at apical portion, widely and deeply depressed between these sinuses, both sides of sinus protruded and forming a pair of elongate median hooks, dorsal surface of median hook with some bristles, dorsal sclerotized part with an articular processe in the middle and a laminate process at the basal portion on each side (Fig. 7).
Female. Body somewhat darker, longer and wider, eyes smaller, antennae thinner and a little shorter than in male. Caudal margin of 8th abdominal sternite with a pair of prominences and a median notch (Figs. 40-50). Claws of legs A type, except for H. takimotoana, which is E type.
Biological Information. Biological information about Hatchina, is extremely limited. Adults were collected on leaves and flowers of various plant species and sometimes at light. In particular, they were frequently found in springtime on newly extending shoots that were colonized by aphids.

Key to the species-groups of Hatchiana in Japan
In providing a key to the species-groups of Hatchiana, the following features were examined: body pubescence; relative length of the 2nd to 3rd antennal segments; shape of pronotum; presence or absence of swelling on the ventral side of femora; conditions of tarsal claws; shape of 8th abdominal sternite in female; length and size of dorsal lobe, ventral lobe and median hook in male genitalia; and shape of aedeagus. As a result, the genus Hatchiana was divided into 3 species-groups as follows.

1(2) Male genitalia with ventral lobe of paramere thick and much shorter than dorsal lobe in lateral view, thick and rounded at apex in ventral view. [Pronotum wider; disk darker; lateral margins rounded and weakly sinuate in front of posterior angle. Femora yellowish brown, darker at apex. Elytra yellowish brown with a longitudinal dark stripe in middle of each elytron. Femora not expanded ventrally]..……………………………….abei species- group
2(1) Male genitalia with ventral lobe of paramere about as long as dorsal lobe, thin, elongate, strongly curved ventrad at apex in lateral view.

3(4) Ventral lobe of paramere less than three times as long as basal width, thick at base, rapidly narrowing apically from middle in ventral view, strongly bent ventrally and hook-shaped at apex. Elytra yellowish brown with longitudinal darker stripe, sometime blackish brown except lateral margin and suture, or entire yellowish brown. Femora yellowish brown, apex darker, sometimes blackish brown except base……heydeni species - group
4(3) Ventral lobe of parameres more than four times as long as basal width, thin and elongate, subparallel-sided or slightly narrowing apically at least on distal half, obtuse at apex in ventral view. Elytra entirely black or blackish brown except latero-ventoral margin yellowish brown. Femora entirely black or blackish brown, sometimes entire brown. [Pronotum entirely reddish yellow or reddish brown.]……….……….……………….ishiharai species - group

The abei species-group
Key to Japanese species of the abei species-group

1(6) Ventral lobe of paramere weakly and gradually narrowing apically from base, with inner margin in a regular arc in ventral view (Figs. 78-80); dorsal fulcrum obtuse, lying at 29~50% of dorsal lobe from base in position (Figs. 82, 83, 84); median hooks thick and short, about 31% of dorsal lobe in length (Figs. 82, 90).

2(3) Dorsal lobe of paramere long, ventral lobe short, less than one-third of dosal lobe in length in lateral view (Fig. 82); median hook short, robust, less than one-third of dorsal lobe in length in lateral view. Female with 8th sternite subtriangularly prominent on each side of median notch at caudal margin (Fig. 40). Antennae with 2nd segment shorter than 3rd, about 83% in male and about 95% in female of 3rd in length. Whitish yellow in general with blackish median stripe on each elytron. Claws of fore and middle legs C types and hind legs B type in male……………………………H. abei (Nakane et Makino, 1981)
3(2) Dorsal lobe of paramere short, ventral lobe short, more than half of the length of dorsal lobe in lateral view (Figs. 83, 84); median hook slender, about half of the length of dorsl lobe in lateral view. Female 8th sternite roundly produced on each side of median notch at caudal margin (Fig. 41, 42). Antennae with 2nd segment much shorter than 3rd, about 72% in male and about 82% in female in length. Brownish in general, with yellowish brown inner and side margins. Claws of fore legs C type, of middle legs A type and of hind legs A type in male.

4(5) Ventral lobe of paramere shorter, about 57% of dorsal lobe in length in lateral view; median hook also shorter, about 47% of dorsal lobe in length in lateral view (Figs. 83, 87)……………………………………H.tsurugisana (Kiriyama, 2000)
5(4) Ventral lobe of paramere longer, about 90% of dorsal lobe in length in lateral view; medain hook longer, about 60% of dorsal lobe in length in lateral view (Figs. 84, 88)………………………… ………………………………………………………H. unzenensis sp.nov.

6(1) Ventral lobe of paramere rapidly narrowing from base to apical third, then parallel-sided to apex, with inner margin almost straight from base to curving point at apical third (Fig. 81); dorsal fulcrum finger-like, lying at about 60% of dorsal lobe from base in position (Fig. 85); dorsal lobe narrower, with deeper and sharper median notch (Fig. 77). Female 8th sternite M-shaped in the middle of caudal margin (fig. 43). Antennae with 2nd segment shorter than 3rd, about 83% in male and about 95% in female in length. Blackish brown in general, with blackish median stripe on elytra, pronotum broadly blackish brown in median area. Claws of fore legs C type, of middle legs A type and of hind legs A type in male………………………………H. osawai (Nakane et Makino, 1981)

Hatchiana abei (Nakane et Makino, 1982), comb. nov.
[Japanese name: Abe-kubiboso-jyokai]
(Figs.17,18,40,51,52,74,78,82,86,90,94,96,97)

Podabrus abei Nakane et Makino in Makino & Nakane, 1982(1981), Rev. Fac. Sci. Kagoshima Univ. (Earth Sci. & Biol.), (14): 59-60, fig. 3 (Mt. Towada, Owani, Aomori Pref., Japan); Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ., 1(2): 1 (N. Honshu); Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1 (Northern Honshu); Imasaka & Yamaya, 1993, Bull. Nagaoka Mun. Sci. Mus. (28): 52, figs. 6-17, 8-17 (Niigata Pref., Awashima Is.).
Podabrus (Podabrus) abei : Takahashi,1998, Kanagawa-chuho, (122): 30 (Honshu: Tohoku, Northern Kanto).
Podabrus (Hatchiana) abei : Takahashi et al.,1999, Bull. Kurashiki Mus. Nat. Hist., (14): 19-20, figs. 3-8, 36, 62 (Aomori Pref.).
Redescription.
Male. Body brownish yellow; temples behind eyes black; scutellum, meso- and metasterna, abdomen except for posterior margin of each sternal segment blackish brown; median portion of clypeus, apical segments of labial and maxillary palpi, apical parts of mandibles, antennae from 3rd to 11th, and apices of 1st and 2nd segments, disk of pronotum, a longitudinal stripe of each elytron in variety, apices of femora and tibiae, tarsi brown to blackish brown, sometimes disk of pronotum and elytra entirely yellowish brown. Body slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles.
Antennae slender, filiform, 2nd segment short, about 0.83 times as long as 3rd; relative lengths of antennal segments from base as follows: 20: 10: 12: 17: 17: 17: 17: 17: 17: 17: 18.
Pronotum about 0.85 times as wide as head, about 1.3 times as wide as long; posterior angles acute and prominent outwards with weak sinus before angle; disk rugose, with a pair of semicircular convexities at base.
Elytra subparallel-sided, about 1.5 times as wide as pronotum, about 2.8 times as long as wide.
Legs slender; femora mostly straight (Fig. 51); claws with triangular process at base, dentation in fore legs C type, middle legs C type and hind legs B type.
Male genitalia almost oval; ventral lobes of parameres thick and short, about 29% as long as dorsal lobe (Fig. 82), rounded at apex (Fig. 78); dorsal lobe subquadrate, longer than wide (Fig. 74); median notch shallow, about 1/7 as deep as length of dorsal lobe; dorsal fulcrum lying at about 29% of dorsal lobe from base in position; median hooks thick and short, about 31% as long as dorsal lobe (Figs. 82, 90).
Length: 10.3-11.8mm, width: 2.7-3.0mm.
Female. Body somewhat darker, longer and wider. Antennae attaining to middle of elytra. Pronotum 1.0 times as wide as head, about 1.4 times as long as wide. Elytra about 1.5 times as wide as pronotum, about 2.6 times as long as wide. Caudal margin of 8th abdominal sternite with a pair of triangular prominences and a subacute and deep median notch, weakly and roundly projected at caudal corners (Fig. 40). Claws with triangular process at base, all claws A types.
Length: 11.2-13.7mm; width: 3.0-3.8mm.

Specimens examined. 33 males & 53 females from the following localites: Honshu : [Aomori Pref.] Yunosawa, Ikarigaseki; Near Shitamae, Kodomari; Kasamasu-toge, Aomori City; [Iwate Pref.] Koyoshibezawa, Mt. Hayachine; Kuzubezawa, Mt. Hayachine; [Akita Pref.] Hachimantai, Iwamisannai, Kawabe; Yagimaguchi, Mt.Chokai; Oofukazawa, Tamagawa; [Yamagata Pref.] Yamadera-Umakata, Yamagata City; Gomizawa, Oku, Oguni; [Miyagi Pref.] Anatozawa, Futakuchi, Sendai City; Futakuchi- keikoku, Akiu; Akiu; Kunimi-pass, Sendai City; Okushinkawa, Hanayama; [Fukushima Pref.] Hinoemata; Eda, Iwaki City; Masuzawa, Tateiwa; [Tochigi Pref.] Mt. Nyoho, Nikko City; Nikko; Nakanosawa- rindo, Fujiwara; [Niigata Pref.] Koage, Asahi; Komanoyu-spa., Yunotani; Mikuni-pass, Yuzawa; Yagisawa, Yuzawa; Mitsumata-kogen, Yuzawa; Renge-spa., Itoigawa City; Udozawa, Itoigawa City; Awashima Is..
Distribution. Japan: Honshu (Northern districts from Aomori to Niigata Prefs.) (Fig. 94). This species is common in northern Japan from low land to mountainous areas. Its distribution is overlapped with that of H. heydeni in the Chubu and Kanto Districts, and with H. hirugatakensis in the Kanto and Tohoku Districts, but this species is more abundant.
Notes. Adults of northern populations are apt to be paler in body coloration and those of southern populations are generally darker with a broader and darker median stripe on elytra.

Hatchiana tsurugisana (Kiriyama, 2000), comb. nov.
[Japanese name: Tsurugi-kubiboso-jyokai]
(Figs.10,19,20,41,53,54,75,79,83,87,91,95,98,99)

Podabrus (Hatchiana) tsurugisanus Kiriyama, 2000, Jpn. J. syst. Ent., 6(2): 251-252,  figs. 3a, 3b, 3c, 4 (Mt. Tsurugi, Tokushima Pref., Japan).
Redescription.
Male. Body brownish yellow; temples behind eyes black; apical portion of clypeus, apical parts of mandibles, antennae except for base and ventral sides of 1st and 2nd segments, disk of pronotum, meso- and metasterna, scutellum, elytra except for latero-ventral side and suture in variety, apex of tarsi, abdomen except for posterior margin of each sternal segment brown to blackish brown.
Body slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles. Antennae slender, filiform, 2nd segment shorter, about 0.67 times as long as 3rd; relative lengths of antennal segments from base as follows: 25: 10: 15: 23: 23: 25: 26: 26: 26: 25: 26.
Pronotum subquadrate (Fig. 10), about 0.87 times as wide as head, about 1.4 times as wide as long; anterior margin weakly bisinuate, posterior margin weakly arcuate and bordered; lateral margins rounded and scarcely angulate in the middle; posterior angles acute and subprominent outwards, with weak sinus in front of angles; disk rugose, with a pair of semicircular convexities at base.
Elytra subparallel-sided, about 1.46 times as wide as pronotum, about 2.63 times as long as wide.
Legs slender; femora mostly straight (Fig. 53); claws of fore legs C type, of middle and hind legs A types.
Male genitalia almost oval; ventral lobes of parameres thick, about 57% as long as dorsal lobe, rounded at apex (Fig. 83); dorsal lobe subquadrate, wider than long (Fig. 75), arcuate apically on each half of posterior margins; median notch shallow, about 11% as deep as length of dorsal lobe; dorsal fulcrum lying at about 47% of dorsal lobe from base in position (Fig. 83); median hooks thin and short, about 47% as long as dorsal lobe, with a few bristles on dorsal surface (Figs. 83, 87, 91).
Length: 11.1-11.8mm, width: 2.6-3.0mm.
Female. Body blackish brown, longer and wider than male; mouth parts except for apices, clypeus before base of antennae, both sides of pronotum, latero-ventral side and suture of elytra, and claws brownish yellow. Antennae attaining to basal 2/5 of elytra. Pronotum as wide as head, about 1.2 times as long as wide. Elytra about 1.4 times as wide as pronotum, about 2.5 times as long as wide. Caudal margin of 8th abdominal sternite narrowly produced caudad and arcuate on each side of shallow meddian notch (Fig. 41). Claws of all legs A type.
Length: 11.2-13.0mm; width: 3.0-3.8mm.

Specimens examined. 22 males & 9 females from following localites: Shikoku: [Kagawa Pref.] Mt. Daisenzan, Kotonami, N. Sano leg.: 3 males, 11.VI.1994, 1 male, 1.VI.1999. [Tokushima Pref.] 1 female, Mt. Nakao-yama, Koyadaira, 4.VI.1998, M. Yoshida leg.; 1 male, Mt. Kotsusan, Yamakawa, 28.V.1997, M. Kako leg.; 1 female. Mt. Kumoso-yama, Kamiyama, 31.V.1998, M. Yoshida leg.; Mt. Takashiroyama, Kisawa: 1 male, 23.VI.1990, A. Watanabe leg.; 1 female, 7.VI.1998, M. Yoshida leg.; Dosu-toge Pass, Kisawa, M. Kako leg.: 1 male & 1 female, 25.VI.1992; 1 male, 23.V.1998; Kamagatani, Kisawa: 1 male, 23.V.1998, M. Kako leg.; 1 male, 5.VI.1999, N. Sano leg.; 1 female, Ochiai-toge Pass, Higashi-iyayama, 9.VII.1995, N. Sano leg.; 1 female, Ochiai-mibuchi-rindo, Higashi- iyayama, 31.V.1999, N. Sano leg.  [Ehime Pref.] 1 male, Jojusha, Mt. Ishizuchiyama, 5.VI. 1994, M. Shiraishi leg.; 1 female, Mt. Ishizuchiyama, Saijyo City, 28.VI.1998, M. Shiraishi leg.; 1 male, Fujinotani〜 Kanpuzan- tunnel, Saijyo City, 6.V.1998, N. Sano leg. ; 2 females, Omogo-kei, Omogo, 13.VI.1998, N. Sano leg..
Distribution. Japan: Shikoku (Kagawa, Tokushima and Ehime Prefs.). This species was collected in early summer from low to high mountains in the northern half of Shikoku (Fig. 95). It has never been collected from lowland, while parapatric congeners, H. heydeni, H.sanoi sp. nov. and H. takimotoana, have been collected from lowland to mountainous areas.
Notes. This species closely resembles H. abei (Nakane et Makino, 1982), but can be separated from it by the following features: erytra with dark stripes, which are more wider in male and blackish brown in female; the ventral lobes of male genitalia about 57% as long as dorsal lobe; dorsal lobe much shorter than that of H. abei and arcuate caudally on each half of posterior margin.

Hatchiana unzenensis Imasaka, sp. nov.
[Japanese name: Unzen-kubiboso-jokai]
(Figs.21,22,42,55,56,76,80,84,88,92,95,100,101)

Podabrus osawai (nec Makino et Nakane, 1982): Imasaka, 1988, Korasana, 26(1): 20, fig. 1-1. Mt. Ohirayama, Beppu City, Ohita Pref., Japan; Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1 (Unzen-kojigoku & Tashirobaru, Shimabara Peninsula, Nagasaki Pref.); Takakura, 1989, Fukuoka-ken no Kochu-so,: 34, 141 (Mt. Hikosan, Mt. Inugatake, Mt. Fukuchisan & Ueno, Akaike, Fukuoka Pref.).
Male. Body brownish yellow; temples behind eyes black; apical portion of clypeus, apical parts of mandibles, antennae except for base and ventral sides of 1st and 2nd segments, disk of pronotum, meso- and metasterna, scutellum, elytra except for latero-ventral side and suture in variety, apex of tarsi, abdomen except for posterior margin of each sternal segments brown to blackish brown.
Body slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles.
Antennae slender, filiform, 2nd segment short, clearly about 0.77 times as long as 3rd; relative lengths of antennal segments from base as follows: 23: 10: 13: 19: 21: 20: 20: 20: 20: 19: 23.
Pronotum subquadrate, about 0.95 times as wide as head, about 1.4 times as wide as long; anterior margin strongly arcuate; posterior margin arcuate and bordered; posterior angles acute, prominent outwards with sinus in front of angles; disk rugose and strongly convex, with a pair of semicircular convexities at base.
Elytra subparallel sided, about 1.33 times as wide as pronotum, about 2.45 times as long as wide, dorsum irregularly and sparsely rugose, smooth at basal part, recognizably costate.
Legs slender; femora mostly straight (Fig. 55); claws of fore legs C type, of middle and hind legs A types.
Male genitalia almost oval; ventral lobe of parameres thick and long, about 90% as long as dorsal lobe, rounded at apex (Figs. 80, 84); dorsal lobe subquadrate (Fig. 76), weakly arcuate apically on each half of posterior margin; median notch shallow, about 20% as deep as length of dorsal lobe; dorsal fulcrum lying at about 50% of dorsal lobe from base in position (Fig. 84); median hooks thick, about 60% as long as dorsal lobe (Figs. 84, 88).
Length: 9.5-11.8mm; width: 2.7-3.2mm.
Female. Body blackish brown, longer and wider than male; mouth parts except for apices, clypeus before base of antennae, both sides of pronotum, narrow latero-ventral side and suture of elytra, tarsi, and claws brownish yellow. Antennae attaining to basal 2/5 of elytra. Pronotum 1.0 time as wide as head, about 1.4 times as long as wide. Elytra about 1.4 times as wide as pronotum, about 2.2 times as long as wide. Caudal margin of 8th abdominal sternite narrowly produced caudad and arcuate on each side of shallow median notch (Fig. 42); Claws of all legs A types.
Length: 10.8-13.7mm; width: 3.2-4.2mm.

Type series. Holotype: male (Deposited in the Entomological Laboratory, Kyushu Univ., Type No.3120), Unzen-hobaru, Obama Town, Nagasaki Pref., Japan, 25.V.1993, S. Imasaka leg. Paratypes: Kyushu: [Nagasaki Pref.] 14 males & 8 females, same data as the holotype; 1 male, Mt. Myokendake, Mt. Unzen, Obama, 14.VI.1994, S. Imasaka leg.; 2 females, Fukkoshi, Mt. Unzen, Obama, 14.VI.1994, S. Imasaka leg.; 3 males, Kojigoku, Obama, 14.V.1986, S. Imasaka leg.; Tashirobaru, Kunimi, S. Imasaka leg.: 2 males & 1 female, 30.V.1988; 1 male & 1 female, 7.V.1991; 1 female, Senbuki, Shimabara City, 7.VI.1978, S. Imasaka leg. [Saga Pref.] Ideno, Mitsuse, S. Imasaka leg.: 1 male, 13.V.1995; 2 males, 30.V.1995; 4 males, 25.V.1999; 2 males & 1 female, Mt. Sefuriyama, Sefuri, 7.VI.1994, S. Imasaka leg. [Fukuoka Pref.] Mt. Shojidake, Mts. Hikosan, M. Noda leg.: 1 male, 25.V.1966; 1 male, 26.V.1968; 1 female, Mt. Gozendake, Yabe, 4.VI.1989, K. Kido leg. [Ohita Pref.] Mt. Kurodake, Shonai: 1 male, 14.VI.1994, Y. Aoki leg.; 1 male & 1 female, 17.V.1998, S. Ogata leg.; 2 females, Oike-lake, Mt. Kurodake, Shonai, 5.VI.1997, S. Imasaka leg.; Mt. Taisenzan, Mts. Kujuzan, K. Aramaki leg.: 2 males, 18.V.1987; 1 male, 1.VI.1987; 2 males, Hoei-rindo, Mt. Katamukiyama, 11.V.1986, T. Miyake leg.; 2 males, Makinoto Pass, Mts. Kujuzan, 5.VI.1997, S. Imasaka leg. [Kumamoto Pref.] 1 female, Senomoto-kogen, Minamioguni, 6.VI.1990, S. Imasaka leg.; 1 male, Kikuchi-keikoku Valley, Kikuchi City, 12.V.1996, S. Imasaka leg.; 1 male, Komagaeri Pass, Yabe, 26.V.1997, S. Imasaka leg.; Shiiya Pass, Yabe: 1 male, 28.V.1997, M. Ebihara leg.; 1 male, 7.VI.1997, S. Imasaka leg.; 1 male, Susogo, Itsukimura, 14.V.1983, I. Ohtsuka leg.; 1 male, Mt. Shiragadake, Uemura, 17.V. 1997, S. Imasaka leg. [Miyazaki Pref.] 1 male, Umaseno, Takachiho, 6.VI.1994, Y. Aoki leg.; 1 female, Sanshudai, Takachiho, 27.VI.1993, A. Nagai leg.; 1 female, Hori, Kitagawa, 20.VI.1995, T. Kinoda leg.; 1 female, Mt. Mukabakiyama, Nobeoka City, 14.V.1967, A. Nagai leg.; Mt. Shiraiwayama, Gokase: 1 male, 3.VI.1976, Y. Yamawaki leg.; 1 female, 29.V.1979, Y. Yamawaki leg.; 3 males, 1.V.1991, A. Nagai leg.; Mt. Koshikidake, Ebino City, A. Nagai leg.: 1 female, 25.VI.1989; 1 male, 5.V.1992, A. Nagai leg.; 1 male, Ebino-kogen, Ebino City, 24.V.1990, A. Nagai leg. [Kagoshima Pref.] 1 male, Mt. Shibisan, Miyanojo, 29.V.1989, Y. Imakiire leg.; 1 male, Ditto, 19.VI.1989, Y. Imakiire leg.; 1 male, Mt. Kurinodake, Kurino Town, 16.V.1988, Y. Imakiire leg.; 1 male, Mt. Kirishimayama, Makizono Town, 23.V.1991, S. Imasaka leg..
Distribution. Japan: Kyushu (All Prefs.). This species was found throughout Kyushu in mountainous areas higher than 400 m above the sea level (Fig.95).
Notes. This new species is extremely close to H. tsurugisana (Kiriyama, 2000), but is distingished from the latter by the following features: ventral lobes of male genitalia long, about 90% as long as dorsal lobe; dorsal lobe longer; the median hooks longer and thicker.
Etymology. The specific name is derived from the name of type locality.

Hatchiana osawai (Nakane et Makino, 1982), comb. nov.
[Japanese name: Osawa-kubiboso-jyokai]
(Figs. 23,24,43,57,58,77,81,85,89,93,94,102,103)
Podabrus osawai Nakane et Makino, in Makino & Nakane, 1982(1981), Rev. Fac. Sci. Kagoshima Univ. (Earth Sci. & Biol.), (14): 61-62, fig.5 (Mt. Ushiroyama, Aida, Okayama Pref., Japan); Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ.,1(2): 1 (Honshu, Kyushu); Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1, (Honshu); Imasaka et al., 1990, Suzumushi, (125): 3, figs. 8-6, 10-6 (Okayama Pref.); Imasaka et Nakamura, 1993, Mis. Rep. Hiwa Mus. Nat. Hist., (31): 48, figs. I-16, III-16 (Hiroshima Pref.); Kiriyama, 1995, Keichitsu, 13(35): 2 (Gifu Pref.).
Podabrus (Podabrus) osawai : Takahashi, 1998, Kanagawa-chuho, (122):30 (Honshu: Chugoku, Kinki, Hokuriku); Kiriyama, 1998, Coleopt. News, (122): 5-6, fig. 2 (Kamikuzukawa, Totsugawa, Nara Pref.).
Redescription.
Male. Body brownish yellow; temples behind eyes black; apical portion of clypeus, apical parts of mandibles, antennae except for base and ventral sides of 1st and 2nd segments, disk of pronotum, meso- and metasterna, scutellum, elytra except for latero-ventral side and suture in variety, apex of tarsi, abdomen except for posterior margin of each sternal segment brown to blackish brown.
Body small and slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles. Antennae slender, filiform, attaining to apical third of elytra; 1st segment clavate, longer and shining; 2nd segment short, about 0.83 times as long as 3rd; relative lengths of antennal segments from base as follows: 20: 10: 12: 19: 19: 19: 19: 19: 19: 18: 20.
Pronotum subquadrate, faintly expanded forwards, about 0.82 times as wide as head, about 1.3 times as wide as long; posterior angles acute, prominent outwards, with sinus before angles; disk rugose and convex such a pair of semicircular convexities at base; medio-longitudinal furrow short and distinct at mddle.
Elytra subparallel sided, about 1.3 times as wide as pronotum, about 2.8 times as long as wide.
Legs slender; femora mostly straight (Fig. 57); claws of fore legs C type, of middle legs and hind legs A types.
Male genitalia almost oval; ventral lobes of parameres a little thin and long, about 75% as long as dorsal lobe, rounded at apex (Figs. 81, 85); dorsal lobe subquadrate, longer than wide (Fig. 77); median notch deep, about 42% as deep as length of dorsal lobe; dorsal fulcrum lying at about 60% of dorsal lobe in position (Fig. 85); median hooks thin, about 63% as long as dorsal lobe (Figs. 85, 89).
Length: 8.0-10.8mm, width: 1.8-2.7mm.
Female. Body somewhat darker, greater part of head, pronotum, elytra and abdomen blackish brown, sometimes lateral side of pronotum reddish brown; antennae attaining to apical half of elytra. Pronotum 1.0 time as wide as head, about 1.2 times as long as wide. Elytra about 1.6 times as wide as pronotum, about 2.5 times as long as wide. Caudal margin of eight abdominal sternite with a M-shaped prominence and a shallow median notch (Fig. 43); latero-caudal angles rounded and weakly produced caudad. Claws of all legs A types.
Length: 9.7-12.7mm; width: 2.3-3.3mm.

Specimens examined. 49 males & 52 females from the following localites: Honshu : [Nagano Pref.] Kuzo, Ohtaki; [Shiga Pref.] Mt. Akagoyama, Yogo; Koiridani Valley, Kutsuki; [Kyoto Pref.] Ashiu, Miyama; Kibune, Kyoto City; Kumogahata, Kyoto City; Mt. Chorogatake; [Mie Pref.] Hirakura, Misugi; [Nara Pref.] Mt. Obakodake; [Wakayama Pref.] Mt. Hoshiyama, Kozagawa; [Hyogo Pref.] Mt. Hyonosen; Akasai-keikoku Valley; [Okayama Pref.] Wakasugi Pass, Nishi-awakura; Takinomiya, Aida; Mt. Kenashiyama, Maniwa-Gun; Ishigoe, Tomana-Gun; Mt. Takiyama, Nagi; [Hiroshima Pref.] Yahata-Kogen, Geihoku; Jyuppozan-rindo, Yoshiwa; Nakatsuya-keikoku Valley, Yoshiwa; [Shimane Pref.] Kabadani, Kakinoki; [Yamaguchi Pref.] Kitanikyo, Nishiki; Notoro, Kawakami Vill.), (Kyushu: [Fukuoka Pref.] 3 males, Mt. Inunakiyama, Mts. Hikosan, 15.V.1997, S. Imasaka leg. New Record).
Distribution. Japan: Honshu (Western districts from Nagano to Yamaguchi Pref.), Kyushu (Fukuoka Pref.) (Fig. 94). This species was collected in low lands to mountain areas in western Honshu and at Mt. Hikosan alone in Kyushu. This is the only species that has distribution pattern overlapped in part with H. unzenensis sp. nov.  in the abei species-group, whereas the other species of the group are allopatric in distribution.
Notes. This species is the smallest among the abei species-group and characterized by having male genitalia with slender ventral lobe and finger-like dorsal fulcrum.

The heydeni species- group
Key to Japanese species of the heydeni species-group

1(2) Dorsal lobe of paramere longer, arcuate on each half of apical margin, median notch clear and sharp (Fig. 119); dorsal fulcrum lying at about 48% of dorsal lobe from base in position (Fig. 125); median hook shorter, about 57% of dorsal lobe in length, narrow at apex, without setae on dorsum (Figs. 128, 131); ventral lobe tapered long to apex (Fig. 122). [Femora weakly swollen internally at a third from apex in both sexes (Figs. 59, 60). Caudal margin of 8th sternite in female produced trapezoidal on each side of median notch (Fig. 44). Pronotum reddish brown to yellowish brown, often infuscate at center. Elytra blackish with yellowish side margin and suture, or sometimes entirely yellowish brown. Claws of fore legs C type, of middle and hing legs B types (Figs. 13, 14)]…………………………………H. heydeni (Kiesenwetter, 1897)
2(1) Dorsal lobe of paramere short in lateral view (Figs. 126, 127), truncate at apical margin, median notch shallow or indistinct (Figs. 120, 121); dorsal fulcurum lying at more than 75% of dorsal lobe from base in position; median hook longer and more than 80% of dorsal lobe in length, thicker, with setae on dorsum (Figs. 126, 127).

3(4) Ventral and dorsal lobes of paramere longer (Figs. 120, 126); median notch of dorsal lobe shallow but distinct (Fig. 120); median hook longer and dorsal setae sparser (Figs. 129, 132). Femora weakly swollen internally at a third from apex in both sexes (Figs. 61, 62). Caudal margin of 8th sternite arcuate caudally on each side of median notch in female. Body slenderer, pronotum yellowish, often infuscate medianly, elytra black with yellowish brown on side margin and suture, sometimes entirely yellowish brown. Claws of fore and middle legs C types, of hind legs B type……………………………H. hirugatakensis (Takahashi, 1992)
4(3) Ventral and dorsal lobes of paramere shorter; dorsal lobe transverse, truncate at apex, with faint or indistinct median notch (Figs. 121, 127); median hook shorter, with denser setae on dorsum (Figs. 130, 133). Body robuster. Femora hardly swollen at a third from apex in both sexes. Caudal margin of 8th sternite weakly produced caudally in middle with shallow median notch in female (Fig. 46). Male pronotum reddish brown, elytra black to blackish brown, with paler suture behind scutellum and side margin. Claws of fore legs C type, of middle and hind legs A types………………………………H. kyushuensis (Takakura, 1988)

Hatchiana heydeni (Kiesenwetter, 1879), comb. nov.
[Japanese name: Kubiboso-jyokai]
(Figs.1-9, 25,26,44,59,60,104,105,119,122,125,128,131,134)

Podabrus heydeni Kiesenwetter, 1879, Dtsch. ent. Z.. 23: 306-307 (Mino, Japan); Lewis, 1895, Ann. Mag. nat. Hist.. Ser.6, 16: 108 (Yuyama, Miyanoshita, Nikko, Suyama); Nakane, 1963, Icon. Ins. Jap. Col. Nat. Edita II: 170, pl. 85-20 (Honshu, Shikoku, Kyushu); Makino & Nakane, 1982(1981), Rev. Fac. Sci. Kagoshima Univ. (Earth Sci. & Biol.), (14): 58-59, fig. 2 (Honshu, Shikoku, Kyushu); Sato, 1985, Col. Japan Color, (III): 108, pl. 17-28 (Honshu, Shikoku, Kyushu); Imasaka, 1988, Korasana, 26(1): 20; Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ., 1(2): 1 (Honshu, Shikoku, Kyushu); Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig. 1 (Honshu, Shikoku, Kyushu); Takakura, 1989, Col. Fauna Fukuoka Pref.: 34-500; Imasaka et all, 1990, Suzumushi, (125): 3, figs. 8-5, 10-5 (Okayama Pref.); Takahashi, 1992, Kanagawa-chuho, (100): 75-76, figs. 1, 47 (Kanagawa Pref.); Imasaka & Nakamura, 1993, Mis. Rep. Hiwa Mus. Nat. Hist., (31): 48, figs. I-15, III-15 (Hiroshima Pref.); Imasaka & Yamaya, 1993, Bull. Nagaoka Mun. Sci. Mus., (28): 52, fig. 6-16 (Niigata Pref.); Kiriyama, 1995, Keichitsu, 13(35): 1-2, (Gifu Pref.); Imasaka & Ohtsuka, 1996, Kumamoto Konchu Doko Kaiho, 40(3): 38-39, fig. 2-1 (Kumamoto Pref.).
Podabrus reinii Heyden, 1879, Dtsch. ent. Z.. 23: 351, (Mino, Japan).
Podabrus heydeni var. immacullicollis Pic, 1905, L'Echange., 21(243): 113 (Japon).
Podabrus (Podabrus) heydeni : Takahashi,1998, Kanagawa-chuho, (122):30 (Honshu, Shikoku, Kyushu); Kiriyama, 1998, Coleopt. News, (122): 5-6, fig. 1 (Kamikuzukawa, Totsugawa, Nara Pref.).
Redescription.
Male. Body brownish yellow; temples behind eyes, meso- and metasterna, scutellum, abdomen except for posterior margin of each sternal segment black; median portion of clypeus, apical segments of labial and maxillary palpi, apical parts of mandibles, antennae from 3rd to 11th segments, longitudinal median stripe of each elytron in variety, apices of femora and tibiae, and tarsi brown to blackish brown, sometimes disk of pronotum, elytra and femora darker, or entirely brownish yellow.
Body slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.77 times as long as 3rd; relative lengths of antennal segments from base as follows: 22: 10: 13: 18: 18: 19: 19: 19: 19: 18: 22.
Pronotum subquadrate (Fig. 9), expanded forwards, about 0.8 times as wide as head, about 1.3 times as wide as long.
Elytra subparallel-sided, about 1.4 times as wide as pronotum, about 2.9 times as long as wide.
Legs slender; femora mostly straight, ventral margin weakly expanded at around 1/3 from apex (Fig. 59); claws of fore and middle legs C types, of hind legs B type.
Male genitalia almost oval; ventral lobes elongate, about 88% as long as dorsal lobe, tapered to apex, bent ventrally (Figs. 122, 125); dorsal lobe subquadrate, wider than long, arcuate on each half of apical margin, with clear and sharp median notch (Fig. 119); dorsal fulcrum lying at about 48% of dorsal lobe from base in position (Fig. 125); median hooks somewhat longer and acute forwards (Fig. 131), about 57% as long as dorsal lobe (Fig. 125), no bristle on dorsal surface( Figs. 128, 131).
Length: 8.6-10.9mm, width: 2.2-2.9mm.
Female. Body somewhat darker; antennae a little shorter than in male, attaining to middle of elytra. Pronotum about 1.1 times as wide as head, about 1.5 times as long as wide. Elytra about 1.6 times as wide pronotum, about 2.7 times as long as wide. Caudal margin of 8th abdominal sternite with a pair of trapezoid prominences in middle and rounded both corners (Fig. 44). Claws of all legs A types.
Length: 10.3-12.8mm, width: 2.8-3.5mm.

Specimens examined. many specimens from the following localites: Honshu : [Gunma Pref.] Kawaba skiing ground, Kawaba; [Niigata Pref.] Mikuni Pass, Yuzawa; [Nagano Pref.] Tobira, Matsumoto City; Mugikusa Pass, Yachiho; Kamikochi; Kuzo, Ohtaki; [Yamanashi Pref.] Gozaishi-kogen; Mt.Ho-o-zan; Enno-machi, Nirasaki City; Mt.Fujisan; [Kanagawa Pref.] Aonohara, Tsukui; [Shizuoka Pref.] Tsurugaike, Iwata City; [Toyama Pref.] Mizunashi-shitsugen, Toga; [Ishikawa Pref.] Nanao City; [Gifu Pref.] Abo Pass; Hiwada-kogen, Takane; Hakusui-lake Shirakawa; Midoridani, Neo; Yaozu Town, Minokamo City; and following Western Prefectures: Aichi, Shiga, Fukui, Kyoto, Mie, Nara, Wakayama, Osaka, Hyogo, Tottori, Okayama, Hiroshima, Shimane, Yamaguchi). Shikoku: Kagawa, Tokushima, Ehime, Kochi. Kyushu: Fukuoka, Saga, Nagasaki, Ohita, Kumamoto, Miyazaki, Kagoshima.
Distribution. Japan: Honshu (Western districts from Gunma to Yamaguchi Prefs.), Shikoku, Kyushu. This species is widely distributed from low to high mountains and from open lands to forests. However, in the northern areas from the Chubu District to North Kanto District, its distribution range is restricted to lower areas and allopatric with the next species, H. hirugatakensis (Takahashi, 1992). However, this species was collected, together with H. hirugatakensis, at high mountainous areas like Kamikochi and Hiwada Plateau around the boundary of Gifu and Nagano Prefs.
Notes. This species is close to H. hirugatakensis, but the body is generally shorter and robuster, the male genitalia are different, and the distribution is allopatric.

Hatchiana hirugatakensis (Takahashi, 1992), comb. nov.
[Japanese name: Hirugatake-kubiboso-jyokai]
(Figs.27,28,45,61,62,106,107,120,123,126,129,132,135)

Podabrus hirugatakensis Takahashi, 1992, Kanagawa-chuho,(100): 92, figs. 2, 48 (Mt. Hirugatake, Tanzawa Mts., Kanagawa Pref., Japan); Takahashi, 1995, Kanagawa-chuho, (113): 20-21, figs. 1−6 (Aomori, Iwate, Fukushima, Tochigi, Yamanashi, Nagano).
Podabrus heydeni (nec Kiesenwetter, 1879): Imasaka & Nakamura, 1993, Mis. Rep. Hiwa Mus. Nat. Hist., (31): 58.
Podabrus (Podabrus) hirugatakensis : Takahashi, 1998, Kanagawa- chuho, (122): 30 (Honshu: from Chubu to Northern district).
Podabrus (Hatchiana) hirugatakensis : Takahashi et al., 1999, Bull. Kurashiki Mus. Nat. Hist., (14): 20-21, figs. 9, 37, 63 (Aomori Pref.).
Redescription.
Male. Body brownish yellow; temples behind eyes, scutellum black; median portion of clypeus, apical segments of labial and maxillary palpi, apical parts of mandibles, antennae from 3rd to 11th and apices of 1st and 2nd segments, meso- and metasterna, abdomen except for posterior margin of each sternal segment, femora, tibiae, and tarsi brown to blackish brown; sometimes disk of pronotum darker, a longitudinal stripe of each elytron in variety darker, or entire brownish yellow.
Body slenderest species in Hatchiana; closely covered with fine yellowish pubescence and fringed with yellowish bristles. Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.77 times as long as 3rd; relative lengths of antennal segments from base as follows: 22: 10: 13: 21: 20: 20: 20: 21: 20: 20: 22.
Pronotum subquadrate, expanded forwards, about 0.82 times as wide as head, about 1.3 times as wide as long; anterior margin weakly arcuate; lateral margins rounded and sinuate.
Elytra subparallel-sided, about 1.5 times as wide as pronotum, about 2.9 times as long as wide.
Legs slender; femora mostly straight, ventral margin weakly expanded at around 1/3 from apex (Fig. 61); claws of fore and middle legs C types, of hind legs B type.
Male genitalia almost oval; ventral lobe almost as long as dorsal lobe, elongate, tapered to apex, but a little shorter than in H. heydeni, bent ventrally (Figs. 123, 126); dorsal lobe subquadrate, wider than long, truncate at posterior margin, median notch clear and shallow (Fig. 120); dorsal fulcrum lying at about 75% of dorsal lobe from base in position (Fig. 126); median hooks thick and long, about 84% as long as dorsal lobe, with a few setae on dorsal surface (Figs. 129, 132).
Length: 9.5-11.2mm, width: 2.3-3.3mm.
Female. Body somewhat darker. Antennae attaining to middle of elytra. Pronotum 1.0 time as wide as head, about 1.5 times as long as wide. Elytra about 1.6 times as wide as pronotum, about 2.7 times as long as wide. Caudal margin of 8th abdominal sternite arcuate caudally on each side of median notch (Fig. 45). Claws of all legs A types.
Length: 12.8-13.5mm, width: 3.0-3.5mm.

Specimens examined. 14 males & 21 females from the following localites: Honshu: [Aomori Pref.] 1 male, Hiratakinuma, Kizukuri, 23.VI.1995, M. Horikawa leg.; [Iwate Pref.] 1 male, Moroe−Fujihichiken, Minami-hachimantai, 25.VII.1979, M. Haga leg.; [Fukushima Pref.] 1 male, Usagidaira, Mts. Azuma, 28.VI.1997, K. Takahashi leg.; 1 male, Komadome-swamp, Showa, 16.VI. 1986, K. Takahashi leg.; 1 male, Funamata-rindo, Hinoemata, 8.VI.1995, M. Horikawa leg.; [Tochigi Pref.] 1 male, Togenochaya, Mt. Nasudake, 5.VI.1995, M. Horikawa leg.; 1 male, Sanno Pass, Okunikko, 16.VI.1994, K. Takahashi leg.; [Kanagawa Pref.] 1 male, Mt. Tanasawanoatama, Mts. Tanzawa, 22.VI.1988, K. Takahashi leg. (paratype); Kamihikawa-rindo, Daibosatsu: 1 male, 18.VII.1982, M. Nishikawa leg.; 1 female, 12.VII.1992, K. Harusawa leg.; Hikawa- rindo, Enzan City, M. Sawai leg.: 1 female, 4.VII.1983; 1 female, 9.VII.1983; 1 female, 27.VI.1984; 1 female, 28.VI.1984; 1 male, 26.VI.1985; 1 male & 1 female, 20.VII.1986; 1 female, 27.VII.1988, K. Masaki leg.; 1 female, Daibosatsu Pass, 23.VII.1984, I. Matoba leg.; 1 female, 24.VII.1984, I. Matoba leg.; 1 female, Himenoyuzawa, Daibosatsu, 30.VI.1997, H. Kamezawa leg.; 1 male, Kiyosato, Mts. Yatsugatake, 4.VIII.1982, K. Mizuno leg.; [Nagano Pref.] 1 male, Shirakoma-ike, Koumi, 3.VII.1994, M. Hinakura leg.; 1 female, Mugikusa Pass, Chino City, 27.VII.1997, H. Kamezawa leg.; 1 male, Tokugo Pass−Iwanadome, Azumi, 15.VII.1993, K. Takahashi leg.; 3 females, Kamikochi 20.VII.1986, S. Takeda leg.; 1 female, Higashiyama, Kiso- fukushima, 20.VII.1986. K. Hosokawa leg.; [Gifu Pref.] 6 females, Hiwada- kogen Takane, 24.VII.1995, H. Kanie leg..
Distribution. Japan: Honshu (Northern Districts from Aomori to Gifu Prefs.). The distribution range of this species partly overlaps with that of H. heydeni in the Chubu Districts, but is generally restricted to higher places than in the latter.
Notes. This species is easily distinguished from H. heydeni by the slender body, anterior position of dorsal fulcurum, and more robust and longer median hook.

Hatchiana kyushuensis (Takakura, 1988), comb. nov.
(Japanese name: Kyushu-kubiboso-jyokai)
(Figs.29,30,46,63,64,108,109a,109b,121,124,127,130,133,135)

Podabrus kyushuensis Takakura, 1988, Kita-Kyushu no Konchu, 35(3): 152, figs. 16-2, A, B, C (Mt. Takasu, Soedacho, Fukuoka Pref., Japan); Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ., 1(2): 1 (Kyushu); Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1 (Mt. Hikosan, Mt. Kujuzan); Nakane, 1992, Kita-Kyushu no Konchu, 39(2): 80 (Mt. Hikosan, Mt. Shojidake).
Podabrus (Podabrus) kyushuensis: Takahashi, 1998, Kanagawa-chuho,(122): 30 (Kyushu).
Redescription.
Male. Body blackish brown; pronotum reddish brown, without dark spot on disk, temples behind eyes, meso- and metasterna, and scutellum black; clypeus before antennal bases, ventral surface of 1st and 2nd segments of antennae, apical parts of mandibles, and posterior margin of each sternal segment of abdomen pale brown.
Body slender, closely covered with fine yellowish pubescence and fringed with yellowish bristles. Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.83 times as long as 3rd; relative lengths of antennal segments from base as follows: 20: 10: 12: 18: 17: 17: 18: 18: 17: 16: 17.
Pronotum subquadrate, about 0.8 times as wide as head, about 1.3 times as wide as long.
Elytra subparallel sided, about 1.5 times as wide as pronotum, about 2.8 times as long as wide.
Legs slender; femora mostly straight (Fig. 65); claws of fore legs C type, of middle and hind legs A types.
Male genitalia almost oval; ventral lobe of paramere about 86% as long as dorsal lobe, sharply tapered apically on apical third (Fig. 124); dorsal lobe subquadrate, much wider than long, truncated at posterior margin, with indefinite or faint median notch (Fig. 121); dorsal fulcrum lying at about 86% of dorsal lobe from the base in position; median hooks thick, about 86% as long as dorsal lobe (Figs. 127, 133), with many bristles on dorsal surface (Fig. 130).
Length: 8.3-11.1mm, width: 2.1-2.7mm.
Female. Body somewhat darker. Antennae attaining to apical half of elytra. Pronotum about 1.05 times as wide as head, about 1.4 times as long as wide. Elytra about 1.4 times as wide as pronotum, about 2.5 times as long as wide. Caudal margin of 8th abdominal sternite with a pair of weakly arcuate prominences (Fig. 46). Claws of all legs A types.
Length: 10.0-13.2mm; width: 2.2-3.7mm.

Specimens examined. 17 males & 43 females from the following localites: Kyushu : [Fukuoka Pref.] Hata-dam, Kitakyushu City; Mt. Joyama, Munakata; Hikosan- shrine, Mt. Hikosan, Soeda; Buzen-bo, Mt. Hikosan, Soeda; Mt. Hikosan, Soeda; Mt. Shojidake, Mts. Hikosan, Soeda; [Nagasaki Pref.] Kinsenji, Takagi Town; Todoroki- fall, Takagi; Kuroki, Ohmura City; [Ohita Pref.] Mt. Takanosuyama, Yamaguni; Mt. Kurodake, Shonai; Daisen-rindo, Mt. Kujuzan; Mt. Kujuzan; Makinoto Pass, Mts. Kujuzan; Fukakusa, Yoshibu, Kuju; Hoei-rindo, Mt. Katamukiyama; Obiragoe, Ogata; Mt. Katamukiyama, Ogata; [Kumamoto Pref.] Shiiya Pass, Yabe; Mt. Shiratoriyama, Izumi; [Miyazaki Pref.] Mt. Goyodake, Hinokage.
Distribution. Japan: northern half of Kyushu (Fukuoka, Nagasaki, Ohita, Kumamoto Prefs., and northern Miyazaki Pref.). This species occurs from low to high mountains.
Notes. This species is distinguished from H. hikosana by having a femur without swelling, and from H. nishidai sp.nov. by having black legs. Specimens from Mts. Kujuzan, Ohita Pref. showed color variations. In addition to the normal type, some specimens have yellowish brown legs, and others have entirely yellowish brown body except the dorsal side of head (Fig. 109b).

The ishiharai species- group
Key to Japanese species of the ishiharai species- group

1(4) Dorsal lobe of paramere longer than wide, truncate at apex, with deep and narrow median notch (Figs. 136, 137); median hook robust, short (Figs. 151, 152), less than 62% as long as dorsal lobe in lateral view, with pointed lateral process. Legs blackish brown in both sexes, femora not swollen at around a third from apex. Caudal margin of 8th sternite in female subtriangularly produced caudad on each side of median notch and latero-caudal corners produced a little longer as round processes (Figs. 47, 48). Antennae with 2nd segment long, 83-86% in male, about 96% in female as long as 3rd.

2(3) Median hook of genitalia shorter, about 33% as long as dorsal lobe in lateral view, with round apex (Figs. 146, 151, 156); dorsal fulcrum close to base in position, lying at about 18% of dorsal lobe in position in lateral view (Fig. 146)….H. ishiharai (M. Sato, 1986)
3(2) Median hook of genitalia longer, about 62% as long as dorsal lobe in lateral view, with squarish apex (Figs. 147, 152, 157); dorsal fulcrum distant from base, lying at about 45% of dorsal lobe in position in lateral view (Fig. 147)………………H. sanoi sp. nov.
4(1) Dorsal lobe of paramere shorter than wide, arcuate on each side of median shallow notch at apical margin (Figs. 138, 139, 140); median hook longer, more than 71% of dorsal lobe in position in lateral view. Femora swollen internally at around a third from apex in both sexes. Caudal margin of 8th sternite in female arcuate caudad on each side of shallow median notch. Antennae with 2nd segment shorter, 71-77% in male, 80-84% in female as long as 3rd.

5(8) Male genitalia with median hooks shorter, less than 79% of dorsal lobe in length in lateral view (Figs. 148, 149); [aedeagus between caudal and subcaudal sinuses widely depressed]; ventral lobe sharply angular at apex in ventral view (Figs. 143, 144). Femora slightly produced in an arch in male (Figs. 69, 70). Claws alike in both sexes, A type.

6(7) Legs yellowish brown in male. Body robuster and shorter; elytra blackish brown, pronotum yellowish brown. Median hook shorter, about 71% as long as dorsal lobe in lateral view (Fig. 148); ventral lobe shorter, about 92% as long as dorsal lobe in lateral view (Fig. 148)……………………………………………………H. nishidai sp. nov.
7(6) Legs black to blackish brown in both sexes. Body slender; elytra darker, pronotum reddish brown; [femora around a third from apex weakly, but distinctly swollen (Figs. 70, 71)]. Median hook longer, about 92% as long as dorsal lobe in lateral view (Fig. 149); ventral lobe longer, about 107% as long as dorsal lobe in lateral view (Fig. 149)……………….H. hikosana (Nakane et Makino, 1981)

8(5) Male genitalia with median hooks longer, a little beyond dorsal lobe at apex in lateral view (Fig. 155); ventral lobe broader, obtuse at apex in ventral veiw (Fig. 145), strongly curved ventrally in lateral view (Fig. 150). Legs black in both sexes, femora obtuse triangularly expanded at a third from apex in male (Fig. 72). Claws different in sexes, all legs B types in male, E types in female (Fig. 16). [Elytra black tinged, pronotum reddish brown. Body slender. Caudal margin of fifth ventrite in female weakly arcuate on each side of shallow median notch. Antennae with 2nd segment shorter, about 71% in male or 80-82% in female as long as 3rd]……………………H. takimotoana (Kiriyama, 2000)

Hatchiana ishiharai (M. Sato,1986), comb. nov.
(Japanese name: Shikoku-kubiboso-jyokai)
(Figs.31,32,47,65,66,110,111,136,141,146,151,156,161)

Podabrus ishiharai M. Sato, 1985, Coleopt. Jpn. Color, 3: 108, pl.17, fig. 29 (Shikoku) (Invalid according to ICZN Art. 13.1.1)
Podabrus ishiharai M.Sato, 1986, Trans. Shikoku ent. Soc.. 17(4): 255-256, figs. 1, 8, 9, 12 (Ishizuchi Skyline, Mts. Ishizuchi, Ehime Pref., Japan); Nakane & Makino, 1989, Rev. Miyazaki Sangyo- keiei Univ., 1(2): 1 (Shikoku); Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1, (Shikoku).
Podabrus (Podabrus) ishiharai : Takahashi,1998, Kanagawa-chuho, (122): 30 (Shikoku).
Redescription.
Male. Body blackish brown; temples behind eyes black; pronotum reddish brown; clypeus before antennal base, basal segments of labial and maxillary palpi, basal parts of mandibles, 1st and 2nd segments of antennae pale brown.
Body slender and small, closely covered with fine yellowish pubescence, antennae and femora with dark-brownish pubescence in addition; apical margin of clypeus, temples and lateral margins of pronotum fringed with yellowish bristles; elytra with yellowish bristles in addition to pubescence.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.86 times as long as 3rd; relative lengths of antennal segments from base as follows: 22: 10: 12: 18: 20: 21: 22: 23: 20: 18: 24.
Pronotum subquadrate, faintly expanded forwards, about 0.87 times as wide as head, about 1.3 times as wide as long; posterior margin arcuate and bordered; lateral margins almost straight; posterior angles subacute and weakly prominent outwards; disk rugose and weakly convex at both side on base.
Elytra subparallel sided, about 1.4 times as wide as pronotum, about 3.0 times as long as wide.
Legs slender; femora straight (Fig. 65); claws of fore legs C type, of middle and hind legs B types.
Male genitalia almost oval; ventral lobes of parameres thin, elongate, as long as dorsal lobe, acute and bent inwards at apex (Fig. 141); dorsal lobe subquadrate, longer than wide, posterior margin truncate (Fig. 136); median notch deep, about 42% as deep as length of dorsal lobe; dorsal fulcrum lying at about 18% of dorsal lobe from base in position (Fig. 146); median hooks very short, about 33% as long as dorsal lobe in lateral view (Figs. 146, 151), rounded at apex, with acute angle at sides and many bristles on basal surface (Fig. 156).
Length: 9.1-9.6mm, width: 2.1-2.3mm.
Female. Body somewhat blackish, except for pronotum reddish brown. Antennae attaining to apical half of elytra. Pronotum about 1.07 times as wide as head, about 1.4 times as long as wide. Elytra about 1.4 times as wide pronotum, about 2.8 times as long as wide. Caudal margin of 8th ventrite with a pair of triangular prominences and an acute and deep median notch, parabolic and more produded than median prominences at latero-caudal corners. Claws of all legs A types.
Length: 10.8-11.9mm; width: 2.7-3.1mm.

Specimens examined. 8 males & 8 females from the following localities: Shikoku: [Ehime Pref.] 1 male, Nibukawa, Tamagawa, 3.V.1996, M. Shiraishi leg.; 2 females, Mt. Saragamine, Shigenobu, 6.VI.1993, N. Sano leg.; Jyojusha, Mt. Ishizuchiyama, Saijyo City, M. Shiraishi leg.: 2 males, 19.VI.1994; 1 female, 26.VI.1994; Mt. Ishizuchiyama, Saijyo City, M. Shiraishi leg.: 2 females, 24.V.1998; 1 female, 31.V.1998; [Kochi Pref.] 5 males & 2 females, Mt. Takanosuyama, Kamegamori-rindo, Hongawa, 6. V.1998, N. Sano leg..
Distribution. Japan: Shikoku (Ehime and Kochi Prefs.). This species was collected only from forests at the Ishizuchiyama mountain range (Fig. 161).
Notes. This species is easily distinguished from parapatric congeners, H. tsurugisana, H. heydeni and others by the blackish and slender body.

Hatchiana sanoi Imasaka, sp. nov.
(Japanese name: Sano-kubiboso-jyokai)
(Figs.33,34,48,67,68,112,113,137,142,147,152,157,161)

Male. Body blackish brown; temples behind eyes black; pronotum reddish brown; clypeus before antennal bases, ventral surface of 1st and 2nd segments of antennae pale brown.
Body slender and small, closely covered with fine yellowish pubescence, antennae and femora admixed with dark-brownish pubescence; apical margin of clypeus, temples and lateral margins of pronotum fringed with yellowish bristles; elytra with yellowish bristles in addition to pubescence.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.83 times as long as 3rd; relative length of antennal segments from base as follows: 20: 10: 12: 18: 18: 19: 19: 20: 19: 18: 18.
Pronotum subquadrate, faintly expanded forwards, about 0.84 times as wide as head, about 1.3 times as wide as long; lateral margins almost straight; posterior angles acute and prominent outwards with weakly sinus before angle.
Elytra subparallel sided, about 1.4 times as wide as pronotum, about 2.9 times as long as wide; dorsum irregularly and sparsely rugose, smooth at basal part, recognizably costate.
Legs slender; femora straight (Fig. 67); tibia feebly bent at base,  mostly straight toward apex; claws of fore legs C type, of middle and hind legs B types.
Male genitalia almost oval; ventral lobes of parameres thin and rather long, about 115% as long as dorsal lobe in lateral view (Fig. 147), acute and incurved at apex in ventral view(Fig. 142); dorsal lobe subquadrate, longer than wide, almost straight at posterior margin, with triangular expansion outwards at side angle, median notch deep, about 45% as deep as length of dorsal lobe (Fig. 137); dorsal fulcrum lying at about 45% of dorsal lobe from base in position (Fig. 147); median hooks about 62% as long as dorsal lobe in lateral view, with some bristles on dorsal surface (Figs. 147, 152), subquadrate at apex with posterior and lateral acute angules in dorsal view (Fig. 157).
Length: 8.5-9.8mm, width: 2.1-2.5mm.
Female. Body somewhat blackened; pronotum reddish brown, longer and wider, eyes smaller and antennae a little shorter than in male; antennae attaining to apical half of elytra. Pronotum about 1.07 times as wide as head, about 1.4 times as long as wide. Elytra about 1.5 times as wide pronotum, about 2.8 times as long as wide. Caudal margin of 8th abdominal sternite with a pair of triangular pointed prominences and a subacute and deep median notch, triangularly produced posteriorly beyond median prominences at side corners (Fig. 48). Claws of all legs A types.
Length: 9.8-12.3mm; width: 2.5-3.6mm.

Type series. Holotype: male (Deposited in the Entomological Laboratory, Kyushu Univ., Type No. 3121), Mt. Takashiroyama, Kisawa, Tokushima Pref., Shikoku, Japan, 1.VI.1986, A. Watanabe leg. Paratypes: Shikoku: [Tokushima Pref.] 1 male, Dosu-toge Pass, Kamiyama, 17.V.1981, M. Yoshida leg.; 4 males, 1 female, Dosu-toge Pass, Kisawa, 4.V.1998, M. Yoshida leg.;1 male & 2 females, same data as the holotype; Mt. Takashiroyama, Kisawa, A. Watanabe leg.: 1 male & 4 females, 15.VI.1986; 2 males & 1 female, 27.V.1990; 2 females, 23.VI.1990; Ditto, M. Yoshida leg.: 1 male, 15.V.1996; 4 females, 31.V.1998; 9 females, 7.VI.1998; 1 female, Kamagatani, Kisawa, 23.V.1998, M. Yoshida leg.; 2 females, Meotoike, nr. Mt. Tsurugisan, Ichiu, 4.VI.1998, M. Kako leg.; 3 females, Ochiai-toge Pass, Higashi-iyayama, 9.VII.1995, N. Sano leg.; 3 females, Iyayama-rindo, Higashi-iyayama, 9.VI.1999, N. Sano leg.; [Kochi Pref.] 1 female, Yahazu-toge Pass, Monobe, 20.VI.2000, N. Sano leg.).
Distribution. Japan: Shikoku (Tokushima and Kochi Prefs.). This species was collected only from the Tsurugisan mountain range, at forests higher than 800 m above the sea level.
Notes. This new species closely resembles H. ishiharai (M. Sato, 1986), but can be distinguished by the following features: dorsal fulcrum of male genitalia locating more distally; median hooks longer; in female, a pair of triangular prominences of the 8th abdominal sternite weaker and shorter. It is also distinguished from the parapatric H. tsurugisana and H. heydeni by the black and slender body. H. takimotoana has coloration same as this species, but the male genitalia, the female claws and the caudal margin of the 8th sternite are different.
Etymology. The specific name was dedicated to Mr. Nobuo Sano (Kotohira Town, Kagawa Pref.), who gave me many specimens and information on the new species.

Hatchiana nishidai Imasaka, sp. nov.
(Japanese name: Tara-kubiboso-jyokai)
(Figs.35,69,114,138,143,148,153,158,161 )

Podabrus ishiharai ssp.? (nec M. Sato, 1986) : Imasaka et Nishida, 1989, Koganemushi, Nagasaki, (50): 4-5, figs.1, 2-7, 7a, 7b (Todoroki-no-taki, Mt. Taradake, Nagasaki Pref., Japan).
Male. Body brownish yellow; temples behind eyes black; clypeus at middle of anterior margin, apical segments of labial and maxillary palpi, apical parts of mandibles, antennae except for ventral side and basal part of 1st segment, meso- and metasterna, scutellum, elytra except for latero-ventral side, tarsi, abdomen except for posterior margin of each sternal segment brown to blackish brown.
Body slender, closely covered with mixture of fine yellowish and darker pubescence; apical margin of clypeus, temples and lateral margins of pronotum fringed with mixture of yellowish and darker bristles; elytra with mixture of yellowish and darker bristles in addition to pubescence.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.77 times as long as 3rd; relative lengths of antennal segments from base as follows: 22: 10: 13: 18: 17: 18: 19: 18: 18: 17: 20.
Pronotum subquadrate, faintly expanded forwards, about 0.88 times as wide as head, about 1.4 times as wide as long; posterior angles subacute and prominent outwards, with weakly sinus in front of angle.
Elytra subparallel sided, about 1.3 times as wide as pronotum, about 2.9 times as long as wide, dorsum irregularly and sparsely rugose, smooth at basal part, recognizably costate.
Legs slender; femora straight, ventral margin weakly expanded at 1/3 portion from apex (Fig. 69); claws of fore legs C type, of middle and hind legs of B types.
Male genitalia almost oval; ventral lobes of parameres thin, elongate, acute and bent inwards at apex in ventral view (Fig. 143), about 92% as long as dorsal lobe in lateral view (Fig. 148); dorsal lobe subquadrate, wider than long, weakly arcuate on each half, with shallow median notch (Fig. 138); dorsal fulcrum lying at about 65% of dorsal lobe from base in position (Fig. 148); median hooks distinctly long, about 71% as long as dorsal lobe (Figs. 148, 153), with a few bristles on dorsal surface, acute and upturned at apex (Figs. 153, 158).
Length: 10.0-10.8mm, width: 2.5-2.6mm.
Female. Unknown.
Type series. Holotype: male (Deposited in the Entomological Laboratory, Kyushu Univ., Type No.3122), Todorokinotaki Fall, Mt. Taradake, Nagasaki Pref., Kyushu, Japan, 6.V.1989, S. Imasaka leg. Paratypes: Kyushu: [Saga Pref.] 1 male, Mt. Kurokamiyama, 10.V.1987, M. Nishida leg.; [Nagasaki Pref.] same locality as the holotype, S. Imasaka leg.: 1 male, 24.V.1994; 1 male, 21.V.2000; 1 male, Taradake-odan-rindo, Takagi, 10.V.1997, N. Yamamoto leg.
Distribution. Japan: Western Kyushu (Saga and Nagasaki Prefs.). Hatchiana nishidai is rare species, and only 5 males were collected in broad leaved evergreen forests 350m〜800m above the sea level on the Taradake mountain range in Nagasaki Pref. and on the summit of Mt. Kurokamiyama (518 m alt.) in Saga Pref.
Notes. This new species is extremely close to H. hikosana (Nakane et Makino, 1982), but distinguishable from it as follows: the male legs are yellowish brown, the expansion at posterior margin of femora is weaker, the elytra are more light-colored, and the ventral lobes of male genitalia and median hooks are shorter.
Etymology. This new species is named in honour of Mr. Mitsuyasu Nishida (Ureshino Town, Saga Pref.), who gave me the specimens and information on locality and emergence period.

Hatchiana hikosana (Nakane et Makino, 1982), comb. nov.
(Japanese name: Hikosan-kubiboso-jyokai)
(Figs.36,37,49,70,71,115,116,139,144,149,154,159,161)
Podabrus hikosanus Nakane et Makino in Makino & Nakane, 1982 (1981), Rev. Fac. Sci. Kagoshima Univ. (Earth Sci. & Biol.), (14): 60-61, fig. 4 (Mt.Hikosan, Fukuoka Pref., Japan); Nakane & Makino, 1989, Rev. Miyazaki Sangyo-keiei Univ., 1(2): 1 (Kyushu); Takakura, 1989, Coleopt. Fauna Fukuoka Pref.,: 34, 266-267, 342 (Mt. Hikosan, Mt. Takanosuyama, Mt. Koshozan); Yamawaki, 1989, Kita-Kyushu no Konchu, 36(2): 137-138; Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig.1, (Kyushu).
Podabrus (Podabrus) hikosanus : Takahashi,1998, Kanagawa-chuho, (122): 30 (Honshu, Kyushu).
Redescription.
Male. Body blackish brown; temples behind eyes black; pronotum reddish yellow; antennae, base of clypeus, basal segments of labial and maxillary palpi, basal parts of mandibles, ventral side of 1st to 3rd segments of antennae, latero-ventral side of elytra, and claws pale brown.
Body slender, closely covered with mixture of yellowish and darker pubescence; apical margin of clypeus, temples and lateral margins of pronotum fringed with mixture of yellowish and darker bristles; elytra with mixture of yellowish and darker bristles in addition to pubescence.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.77 times as long as 3rd; relative lengths of antennal segments from base as follows: 22: 10: 13: 18: 19: 19: 19: 18: 18: 17: 18.
Pronotum subquadrate, faintly expanded forwards, about 0.87 times as wide as head, about 1.3 times as wide as long.
Elytra subparallel sides, about 1.4 times as wide as pronotum, about 2.9 times as long as wide.
Legs slender; femora straight, ventral margin strongly and roundly expanded at around 1/3 from apex (Fig. 70); claws of fore legs C type, of middle and hind legs B types.
Male genitalia almost oval; ventral lobes of parameres thin and elongate, acute at apex and incurved in ventral view (Fig. 144), about 107% as long as dorsal lobe (Fig. 149); dorsal lobe subquadrate, wider than long; posterior margin weakly arcuate on each half, with shallow median notch (Fig. 139); dorsal fulcrum lying at about 67% of dorsal lobe from base in position (Fig. 149); median hooks distinctly longer, about 79% as long as dorsal lobe (Figs. 149, 154), with many bristles on dorsal surface, acute and upturned at apex (Figs. 154, 159).
Length: 8.0-10.8mm, width: 1.8-2.7mm.
Female. Body black, pronotum reddish brown, mouth parts, vental sides of antennal base, latero-ventral side of elytra, and claws pale brown. Antennae attaining to apical half of elytra. Pronotum about 1.05 times as wide as head, about 1.35 times as long as wide. Elytra about 1.4 times as wide pronotum, about 2.7 times as long as wide. Caudal margin of 8th abdominal sternite with a pair of arcuate central prominences on each subacute side and fairly deep median notch, rounded at latero-caudal corners. Femora strongly and roundly expanded posteriorly at around 1/3 from apex (Fig. 71); Claws of all legs A types.
Length: 9.7-12.7mm; width: 2.3-3.3mm.

Specimens examined. 10 males and 9 females from the following localities. Kyushu : [Fukuoka Pref.] Mt. Jyoyama, Munakata, 1 female, 5.VI.1983, K. Kido leg.; 1 female, 1.VI.1984, S. Ogata leg.; 1 male & 1 female, 10.V.1998, S. Imasaka leg.; 1 female, 18.V.1998, S. Imasaka leg.; Mt. Kasagiyama, Iizuka City, Y. Yamawaki leg.: 1 male, 24.V.1988; 5 males, 30.V.1988; 1 female, 4.VI.1988; 1 male, 23.V.1990; 2 females, 12.V.1992; 1 male, Hikosan, Buzen, Kyushu, 8.VI.1937, Esaki & Hori leg.(Collection of Kyushu Univ.); [Ohita Pref.] 1 female, Karimata-yama, Yamaguni, 10.V.1997, F. Hirokawa leg..
Distribution. Japan: Northern Kyushu (Fukuoka and Ohita Prefs.)
Notes. This species is easily distinguished from H. unzenensis sp. nov., H. osawai and H. heydeni by the reddish pronotum and black and slender body, and from H. kyushuensis by the swollen femora.

Hatchiana takimotoana (Kiriyama, 2000), comb. nov.
(Japanese name: Takimoto-kubiboso-jyokai)
(Figs.11,38,39,50,72,73,117,118,140,145,150,155,160,162)
Podabrus (Hatchiana) takimotoanus Kiriyama, 2000, Jpn. J. syst. Ent., 6(2): 249-251,  figs. 1a, 1b, 1c, 2 (Takimoto, Kumanogawa-cho, Wakayama Pref., Japan).
Podabrus hikosanus ssp.? (nec Makino et Nakane, 1982): Imasaka & Nishida, 1989, Koganemushi, (50): 4-5, fig. 1 (Hiroshima, Okayama); Imasaka et al., 1990, Suzumushi, (125): 3, figs. 8-7, 10-7 (Mt. Hoki-daisen; Yoshiwa, Hiroshima Pref.; Okayama Pref.); Imasaka & Nakamura, 1993, Misc. Rep. Hiwa Mus. nat. Hist.., (32): 49, figs. I-17, III-17 (Hiroshima).
Podabrus sp.: Mizota & Imasaka, 1997, Bull. Hokkaido Univ. For., 54(2): 320 (Kozagawa, Wakayama Pref.).
Redescription.
Male. Body blackish brown; temples behind eyes black; pronotum reddish yellow; Apices of clypeus, mandibles, 1st to 2nd segments of antennae, and claws brown; sometimes latero-ventral margins of elytra nallowly pale brown.
Body very slender and larger in size, closely covered with yellowish and darker pubescence; apical margin of clypeus, temples and lateral margins of pronotum fringed with yellowish and darker bristles; elytra with yellowish and darker bristles in addition to pubescence.
Antennae slender, filiform, attaining to apical third of elytra; 2nd segment about 0.71 times as long as 3rd; relative lengths of antennal segments from base as follows: 24: 10: 14: 19: 19: 20: 22: 22: 21: 19: 21.
Pronotum subquadrate (Fig. 11), faintly expanded in the middle, about 0.95 times as wide as head, about 1.3 times as wide as long; posterior margin strongly arcuate and bordered; lateral margins sinuate; anterior angles fairly angulate; posterior angles strongly acute and prominent outwards, with sinus in front of angle.
Scutellum triangular.
Elytra subparallel sided, about 1.4 times as wide as pronotum, about 2.9 times as long as wide.
Legs slender; femora straight, strongly and triangularly expanded ventrally at around 1/3 from apex (Fig. 72); claws of fore legs D type, of middle and hind legs B types.
Male genitalia almost oval; ventral lobes of parameres thick, long and weakly incurved in ventral view (Fig. 145), about 108% as long as dorsal lobe (Fig. 150), strongly curved ventrally in latral view (Fig. 150); dorsal lobe subquadrate, wider than long, weakly arcuate on each half and shallow notch at middle of posterior margin, angulate at lateral corners (Fig. 140); dorsal fulcrum lying at about 86% of dorsal lobe from base in position (Fig. 150); median hooks especially long, about 1.04% as long as dorsal lobe (Figs. 150, 160), with a few bristles on dorsal surface, with acute and upturned apex (Figs. 155, 160).
Length: 10.7-12.8mm, width: 2.5-3.0mm.
Female. Body somewhat black with almost blackish brown pubescence and bristles; pronotum reddish yellow. Antennae a little shorter than in male, the latter attaining to apical half of elytra. Pronotum about 1.15 times as wide as head, about 1.5 times as long as wide. Elytra about 1.4 times as wide as pronotum, about 2.8 times as long as wide. Caudal margin of 8th sternite with a sinuate prominence and a shallow median notch, rounded on both corners. Femora strongly and roundly expanded ventrally at around 1/3 from apex (Fig. 73); Claws of legs peculiar in form, with short tooth and triangular process at base, all of then E types.
Length: 12.5-14.7mm; width: 3.3-3.8mm.

Specimens examined. 30 males and 26 females from the following localities. Honshu : [Nara Pref.] 1 female, Mt. Obakodake, Nosegawa, 21.VI.1987, T. Hatayama leg.; 1 female, Ohmata, Mt. Obakodake, Nosegawa, 9.VII.1994, T. Hatayama leg.; [Wakayama Pref.] 1 female, Osugi-zuido, Hongu, 11.VI.1994, K. Harusawa leg., 1 male & 6 females, Mt. Hoshiyama, Kozagawa, 7.VI.1995, K. Mizota leg.; [Tottor Pref.] Mt. Hoki-daisen: 1 male, 13.VI.1971, S. Imasaka leg.; 1 female, 14.VI.1986, H. Okada leg.; 1 male, 21.VI.1987, H. Okada leg.; 1 female, Mt. Karasugasen, Mts. Hoki-daisen, 17.VI.1995, F. Hirokawa leg.; [Okayama Pref.] 1 male, Sangajyo, Kamisaibara, 26.VI.1971, O. Yamaji leg.; 1 male, Mt. Gagyuzan, Takahashi City, 22.V.1971, O. Yamaji leg.; [Hiroshima Pref.] Nakatsuya-keikoku Valley, Yoshiwa, S. Imasaka leg.: 1 male, 23.V.1988; 1 male, 24.V.1988; Ditto, T. Nakahara leg.: 1 male, 5.VI.1988; 1 female, 25.VI.1988; 1 female, Mt. Gori, 9.VI.1979, H. Aramaki leg. Shikoku : [Kagawa Pref.] 11 males & 1 female, Mt. Daisenzan, Kotonami, Kagawa Pref., 11.VI.1994, N. Sano leg.; 1 male, Mt. Kasagata, Kotonami, 22.V.1989, N. Sano leg.; 1 female, Kawaoku, Kotonami, 7.VI.1998, N. Sano leg.; Mt. Zozusan, Kotohira, N. Sano leg.: 1 male, 13.V.1994; 1 female, 28.V.1994, N. Sano leg.; [Tokushima Pref.] Mt. Ryuozan, Mima, N. Sano leg.: 1 female, 15.VI.1998; 1 female, 19.VI.2000; 1 male & 1 female, Mt. Otakiyama, Wakimachi, 13.VI.1976, M. Yoshida leg.; Mt. Takashiroyama, Kisawa: 1 female, 26.VI.1991, A. Watanabe leg.; 1 male, 20.VII.1991, M. Kako leg.; Dosu-toge Pass, Kisawa, M. Kako leg.: 1 female, 25.VI.1992; 2 males & 1 female, 23.V.1998; Ditto, 1 female, 31.V.1998, M. Yoshida leg.; 1 male, Kuwadaira, Ichiu, 14.VII.1991, M. Kako leg.; 1 male, Mt. Tonomaru, Ichiu, 4.VI.1998, M. Yoshida leg.; 2 females, Mt. Tsurugisan, 16-17.VII.1994, K. Masaki leg.; 1 male, Mt.Kumosoyama, Kamiyama, 31.V.1998, M. Yoshida leg.; 1 male & 1 female, Mitsugi, Kamiyama, 31.V.1998, M. Yoshida leg.; [Ehime Pref.] 1 male, Mt. Ishizuchiyama, 31.V.1998, M. Shiraishi leg.).
Distribution. Japan: Honshu (Nara, Wakayama, Tottori, Okayama, and Yamaguchi Prefs.), Shikoku (Kagawa, Tokushima, and Ehime Prefs.). This species was collected mainly in the Fagus vegetation zone.
Notes. This species closely resembles H. hikosana (Nakane et Makino, 1982), but differs from it as follows: body more robust and slenderer; femora more strongly expanded; claws peculiar in both sexes, especially in female; median prominences of the 8th sternite in female at caudal margin weaker; median hooks of male genitalia longer than dorsal lobe.
Remarks on the distribution pattern of Hatchiana
The genus Hatchiana has never been found in Hokkaido and most of the surrounding islands of Japan including the Ryukyus. The abei and heydeni species-groups are widely distributed in Honshu, Shikoku, and Kyushu, but the ishiharai species-group is confined to the montainous areas west of the Kinki Districts. There is a tendency for the number of species to increase towards south-western Japan, for instance 6 species in Kyushu: 5 in Mt. Hiko in Fukuoka Pref., 4 in Mt. Fukuchi in Fukuoka Pref., and 5 species in Shikoku: Mt. Ishizuchi in Ehime Pref., and at most 3 species in the other areas in Honshu.

Acknowledgements
I wish to express my cordial thanks to Prof. Emeritus K. Morimoto of Kyushu Univ., for his comments on the descriptions and encouragements.
Special thanks are due to Prof. J. Yukawa and Assoc. Prof. O. Tadauchi of Kyushu Univ. for their critical reading of early draft, and to the late Dr. T. Nakane, Prof. M. Sato of Nagoya Women's Univ., and Mr. K. Takahashi of Hiratsuka City, for their guidance and encouragements.
My thanks are due to Prof. Emeritus T. Kumata of Hokkaido Univ., Dr. Sadanari Hisamatsu of Matsuyama City, Assist. Prof. S. Kamitani, Mr. T. Ueno, Dr. H. Kojima and Mr. N. Takahashi of Kyushu Univ., Dr. K. Yahiro of Biwako Shiga Pref. Mus., Mr. Y. Okushima of Kurashiki Mus. of Nat. Hist., Mr. K. Mizota of Hokkaido Univ., for providing me with valuable specimens and literature.
I also thank to the following persons for allowing me to examine specimens: Messrs. N. Abiru (Nagasaki City), M. Noda (Nagasaki City),  T. Hatayama (Osaka City), M. Kako (Tokushima City), Dr. Y. Kusui (Kaizuka City), K. Masaki (Kyoto City), Y. Matsunaga (Kitakyushu City), H. Ohishi (Kyoto City), the late T. Nakahara, M. Nishida (Ureshino Town), Dr. S. Nakamura (Nat. Hist. Museum of Hiwa), I. Ohtsuka (Kumamoto City), H. Okada(Akashi City), N. Sano (Kotohira Town), S. Takeda (Takatsuki City), A. Watanabe (Kurashiki City), O. Yamaji (Okayama City), S. Yamaya (Nagaoka Museum of Nat. Hist.), N. Yasuda (Sounkyo Museum of Nat. Hist.), and M. Yoshida (Tokushima City).

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Explanation of Plates
Plate 1. Habitus, head, male genitalia, pronotum and claws of Hatchiana spp.
Fig. 1, habitus; 2, head, showing tentorium in dorsal view; 3, head, ventral view; 4-6, male genitalia (4, dorsal; 5, ventral. 6, lateral); 7-8, aedeagus (median lobe) of male genitalia (7, dorsal; 8, lateral); 9-11, pronotum, dorsal view; 12-16, types of claws (12, A type; 13, B type; 14, C type; 15, D type; 16, E type). 1-9, H. heydeni ; 10, H. tsurugisana ; 11, H. takimotoana.
Abbreviations denoting the term used in description:
a=aedeagus, aat=anterior arm of tentorium, ao=apical orifice, ap=articular process, a1=1st segment of antenna,
a2=2nd segment of antenna, a3=3rd segment of antenna,
bo=basal orifice, bp=basal piece, cl=clypeus, cs=caudal sinus, df=dorsal fulcrum, dl=dorsal lobe, dp=dorsal plate,
dsp=dorsal sclerotized part, e=eye, en=elliptical nodule,
es=ear stand of anterior bridge of tentorium, g=gula,
gs=gular suture, li=lateral incisure, ls=lateral sclerite, lp=laminate process, m=mentum, ma=mandible, mn=median notch, mh=median hook, o=occiput, of=occipital foramen, p=parameres, pat=posterior arm of tentorium, ptp=posterior tentrial pit, s=section of mandible, scs=subcaudal sinus, vl=ventral lobe, vmp=ventral membranous part, vp=ventral plate.
Plate 2. Figs. 17-39, First to 3rd segments of antenna in Hatchiana spp. 40-50, Caudal margin of eight abdominal sternite in female of Hatchiana spp.
17, male, 18 & 40, female: H. abei ; 19, male, 20 & 41, female: H. tsurugisana ; 21, male, 22 & 42, female: H. unzenensis ; 23, male, 24 & 43, female: H. osawai ; 25, male, 26 & 44 female: H. heydeni ; 27, male, 28 & 45, female: H. hirugatakensis ; 29, male, 30 & 46, female: H. kyushuensis ; 31, male, 32 & 47, female: H. ishiharai ; 33, male, 34 & 48, female: H. sanoi ; 35, male: H. nishidai ; 36, male, 37 & 49, female: H. hikosana ; 38, male, 39 & 50, female: H. takimotoana .
Plate 3. Figs. 51-73, Fore, middle and hind femora of Hatchiana spp.
51, male, 52, female: H. abei ; 53, male, 54, female: H. tsurugisana ; 55, male, 56, female: H. unzenensis ; 57, male, 58, female: H. osawai ; 59, male, 60, female: H. heydeni ; 61, male, 62, female: H. hirugatakensis ; 63, male, 64, female: H. kyushuensis ; 65, male, 66, female: H. ishiharai ; 67, male, 68, female: H. sanoi ; 69, male: H. nishidai ; 70, male, 71, female: H. hikosana ; 72, male,73, female: H. takimotoana .
Plate 4. Figs. 74-85, Male genitalia of the abei species-group of Hatchiana.
74-77, dorsal plate, dorsal view; 78-81, ventral plate and apical part of median hook, ventral view; 82-85, dorsal lobe, ventral lobe and median hook, lateral view; 86-89, aedeagus, dorsal view; 90-93, aedeagus lateral view; 74, 78, 82, 86, 90: H. abei ; 75, 79, 83, 87, 91: H. tsurugisana ; 76, 80, 84, 88, 92: H. unzenensis ; 77, 81, 85, 89, 93: H. osawai .
Plate 5. Figs. 94-95, Distributions of the abei species-group of Hatchiana spp.
Fig. 94, ●: H. abei, ○: H. osawai ; Fig. 95, ●: H. tsurugisana, ○: H. unzenensis .
Plate 6. Figs. 96-118. Photographs of Hatchiana spp.
96, male, 97, female: H. abei ; 98, male, 99, female: H. tsurugisana ; 100, male, 101, female: H. unzenensis ; 102, male, 103, female: H. osawai ; 104, male, 10, female: H. heydeni ; 106, male, 107, female: H. hirugatakensis ; 108, male, 109, female, a: normal type, b: brownish type: H. kyushuensis ;  110, male, 111, female: H. ishiharai ; 112, male, 113, female: H. sanoi ; 114, male: H. nishidai ; 115, male, 116, female: H. hikosana ; 117, male, 118, female: H. takimotoana.
Plate 7. Figs. 119-127. Male genitalia of the heydeni species-group of Hatchiana.
119-121, Dorsal plate, dorsal view; 122-124, ventral plate and median hook, ventral view; 125-127, dorsal lobe, ventral lobe and median hook, lateral view; 128-130, aedeagus, dorsal view; 131-133, aedeagus, lateral view; 119,122,125,128,131: H. heydeni ; 120,123,126,129,132: H. hirugatakensis ; 121,124,127,130,133: H. kyushuensis.
Plate 8. Figs. 134, 135. Distributions of the heydeni species-group of Hatchiana.
134, ●: H. heydeni ; 135, ●: H. hirugatakensis ; ○: H. kyushuensis.
Plate 9. Figs. 136-160. Male genitalia of the ishiharai species-group in Hatchiana.
136-140, Dorsal plate, dorsal views; 141-145, ventral plate, ventral view; 146-150, dorsal and ventral lobes, and median hooks, lateral view; 151-160, aedeagus (median lobe), lateral and dorsal views.
136, 141, 146, 151, 156: H. ishiharai ;  137, 142, 147, 152, 157: H. sanoi ; 138, 143, 148, 153, 158: H. nishidai ;  139, 144, 149, 154, 159: H. hikosana; 140, 145, 150, 155, 160: H. takimotoana.
Plate 10. Figs. 161, 162:  Distribution maps of the ishiharai species-group of Hatchiana.

161. ○: H .ishiharai, ●: H. sanoi, □: H. nishidai,  ■: H. hikosana, 162. ●: H. takimotoana.

 


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